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International Journal of Primatology

The Official Journal of the International Primatological Society

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International Journal of Primatology - Visit our Gallery with our stunning cover designs

Welcome to the captivating world of primates showcased through the vibrant covers of the International Journal of Primatology. As stewards of scientific inquiry into the lives and behaviors of our closest evolutionary relatives, we are delighted to present a gallery that celebrates the diversity, complexity, and beauty of the primate realm.

Each cover tells a story—a visual narrative of the fascinating research, discoveries, and conservation efforts that shape our understanding of primates' ecological, social, and cognitive lives. From the dense rainforests of the Amazon to the vast savannas of Africa, and beyond, these covers offer glimpses into the rich tapestry of primate habitats and behaviors across the globe.

Whether it's the striking gaze of a chimpanzee, the intricate grooming patterns of a group of monkeys, or the playful antics of a leaping lemur, each image encapsulates the essence of these remarkable creatures. Through the lens of science and artistry, we invite you to embark on a journey of exploration, appreciation, and understanding of our primate cousins.

Join us as we celebrate the beauty and diversity of primates, and delve deeper into the intricacies of their world, one cover at a time.

February 2024, Volume 45, Issue 1
10764_45_1_CoverFigure_PrintThe image shows a long-calling titi monkey (Plecturocebus modestus). Like the special issue on them within, this is used to summon attention and inform on status. Photo credit: Rob Wallace.
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December 2023, Volume 44, Issue 6
10764_44_6_CoverFigure_PrintThis image shows a bonnet macaque infant (Macaca radiata) and her mother who is grooming another female. The grieving behavior of female bonnet macaques after the loss of their infants is described in this issue by Arlet et al. on pages 1182–1199. Photo by Thenmala Macaque Project. Right-click here to enlarge the image



October 2023, Volume 44, Issue 5
10764_44_5_CoverFigure_PrintA photo of a golden monkey (Cercopithecus mitis kandti) group foraging on potatoes in a potato field directly adjacent to the Volcanoes National Park, Rwanda. This image represents the conflict between farmers and golden monkeys in the form of the latter exiting their natural habitat to forage on potato crops; the effects of the conflict and mitigation measures are studied in this issue by Ndayishimiye et al. on pages 856– 880. Photo by Dian Fossey Gorilla Fund and Eric Ndayishimiye.
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August 2023, Volume 44, Issue 4
10764_44_4_CoverFigure_PrintRelaxed grooming session of a group of crested macaques in Tangkoko Reserve, North Sulawesi, Indonesia. This Special Issue focuses on the behaviour, ecology, and conservation of this critically endangered species, and celebrates the work of the Macaca Nigra Project, which studies the biology of the crested macaques in their natural habitat and promotes its conservation since 2006. Credit: Jérôme Micheletta/the Macaca Nigra Project.
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June 2023, Volume 44, Issue 3
10764_44_3_CoverFigure_PrintIllustration of affiliative interaction between female Japanese macaques (Macaca fuscata). Female Japanese macaques that perform affiliative interactions with each other are studied in this issue by Ishizuka and Inoue on pages 500-516. Credit: Shintaro Ishizuka.
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April 2023, Volume 44, Issue 2
10764_44_2_CoverFigure_PrintThis image depicts a fat-tailed dwarf lemur (Cheirogaleus medius) enjoying a sugary fruit: in this issue, Blanco and Greene on pages 254-257 ask if the seasonal preference for sugar in dwarf lemurs at the Duke Lemur Center matches seasonal diets in the wild. Credit: David Haring. Right-click here to enlarge the image



February 2023, Volume 44, Issue 1
10764_44_1_CoverFigure_PrintThis image represents the posture of the mongoose lemur (Eulemur mongoz) in Mohéli Island when he sees a photographer. The individual was feeding in a clove tree before being drawn into the camera.
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December 2022, Volume 43, Issue 6
10764_43_6_CoverFigure_PrintThis image is an example of the community inclusion approach of Barbary Macaque (Macaca sylvanus) Awareness and Conservation in the Rif Mountains of Morocco described in this issue by Waters et al. on pages 989– 999. Credit: Ahmed El Harrad. Right-click here to enlarge the image



October 2022, Volume 43, Issue 5
10764_43_5_CoverFigure_PrintThis image represents the presence of the golden monkey (Cercopithecus mitis kandti), also known as the golden guenon in the restored forest of the Gishwati-Mukura landscape, which is studied in this issue by Tuyisingize et al. on pages 867– 884. Credit by Deogratias Tuyisingize.
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August 2022, Volume 43, Issue 4
10764_43_4_CoverFigure_PrintThis image of the golden-brown mouse lemur (Microcebus ravelobensis) was taken in Jardin Botanique A, a long-term research site in the Ankarafantsika National Park, northwestern Madagascar, from where this species was first described by Elke Zimmermann and her students in 1998. Photo by Ute Radespiel.
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June 2022, Volume 43, Issue 3
10764_43_3_CoverFigure_PrintThis image shows a fork-marked lemur (Phaner pallescens) during its typical gum-foraging behavior on a tree of the genus Terminalia in Kirindy Forest, Madagascar. Estimating density and vocal activity of Phaner pallescens using passive acoustic recorders is studied in this issue by Markolf et al. on pages 409– 433. Photo by Hanitra Rakotonirina.
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April 2022, Volume 43, Issue 2
10764_43_2_CoverFigure_PrintThis photo was taken in the Pacaya Samiria Reserve of the Peruvian Amazon. A large-headed capuchin (Sapajus macrocephalus) is feeding on urucuri (shapaja) palm fruit (Attalea phalerata) whilst looking at a human observer. Observer effects are studied in this issue by Metcalfe et al. on pages 216 –234. This photograph is copyright of Chloë Alexia Metcalfe. Right-click here to enlarge the image


February 2022, Volume 43, Issue 1
10764_43_1_CoverFigure_PrintLogging and road construction along the boundary of Monte Alen National Park, Equatorial Guinea. The park is home to 16 primate species, including central chimpanzees (Pan troglodytes troglodytes) and western lowland gorillas (Gorilla gorilla gorilla). Photo courtesy of Patrick McLaughlin/Bristol Zoological Society.
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December 2021, Volume 42, Issue 6
10764_42_6_CoverFigure_PrintThis image represents an adult female chimpanzees (Pan troglodytes) of the studied group in ZooParc de Beauval, France. The calling patterns of the vocal interactions within this group are studied in this issue by Pougnault et al. on pages 809–832. Photo by Corinne Pellegrino.
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October 2021, Volume 42, Issue 5
10764_42_5_CoverFigure_PrintA mother and two infant snub-nosed monkeys huddling during a snow storm in Shennongjia National Park, China. The thermoregulatory challenges that snub-nosed monkeys have to face in winter, the endocrine mechanisms of which are studied in this issue by Chen et al. on pages 759–763. Photograph by Zuofu Xiang. Right-click here to enlarge the image


August 2021, Volume 42, Issue 4
10764_42_4_CoverFigure_PrintThis photograph shows a captive Red-mantled Saddle-back tamarin (Leontocebus lagonotus) that is a victim of wildlife trafficking at Yanacocha Wildlife Rescue Center, Puyo, Ecuador. Photography by Diego G. Tirira.
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June 2021, Volume 42, Issue 3
10764_42_3_CoverFigure_PrintThis image is of a juvenile male aye-aye (Daubentonia madagascariensis) clinging to trunk who was part of a study on behavioral development, as described in this issue by Rakotondrazandry et al. on pages 344–348. Photo by Richard Randriamampionona.
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April 2021, Volume 42, Issue 2
10764_42_2_CoverFigure_PrintThis image represents one of the Barbary macaques (Macaca sylvanus) on a snowy day housed in Parco Natura Viva – Garda Zoological Park (Italy). The zoo colony of this macaque was involved in the study by Baldachini et al. on pages 155–170, investigating hand preference for inanimate and animate targets in this species. Photo by Giorgio Ottolini.
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February 2021, Volume 42, Issue 1
10764_42_1_CoverFigure_PrintPhoto of an adult female of the species Aotus lemurinus from the municipality of Pijao, Colombia. This individual was part of the study of the ecology of night monkeys included in this issue by Montilla et al. on pages 130 –153. Photo by Sebastián O. Montilla.
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December 2020, Volume 41, Issue 6
10764_41_6_CoverFigure_PrintMale savanna chimpanzee (Pan troglodytes) sitting in the miombo woodland vegetation at Issa, western Tanzania. Effects of method and season on the party sizes of Issa chimpanzees were studied in this issue by Vink et al. on pages 901– 915. Photo credit attributed to Carolyn Fryns/GMERC.
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October 2020, Volume 41, Issue 5
10764_41_5_CoverFigure_PrintPhoto of Galagoides thomasi with very visible eye shine taken in a small village of Woteva in Southwest Cameroon in 2017. It is one of the nocturnal primates studied in this issue by Fominka et al. on pages 714 –731. Photo by Nestoral Fominka.
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August 2020, Volume 41, Issue 4
10764_41_4_CoverFigure_PrintFemale pileated gibbon (Hylobates pileatus) resting in a tree at Phnom Tamao Wildlife Rescue Center, Cambodia. This pileated gibbon was screened for Hepatitis B virus (HBV). HBV prevalence in wild-living gibbons was studied in this issue by Leroux et al. on pages 634 – 653 . Photo by Jeremy Holden/ Wildlife Alliance.
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June 2020, Volume 41, Issue 3
10764_41_3_CoverFigure_PrintThis image represents a typical quadrupedal sit position used by arboreal Malabar slender lorises (Loris lydekkerianus malabaricus) to sit on vertical terminal branches. The ecological determinants of Malabar slender loris occurrence are studied in this issue by Gnanaolivu et al. on pages 511– 524. Photo by Daniel Divyakumar H.
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April 2020, Volume 41, Issue 2
10764_41_2_CoverFigure_PrintThis image is of a peaceful encounter of three groups of wild bonobos at Wamba, the Democratic Republic of the Congo. Females and their offspring gathered and groomed with one another in the middle on a fallen tree, while adult males were tense and remained at the periphery. Photograph by Takumasa Yokoyama.
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February 2020, Volume 41, Issue 1
10764_41_1_CoverFigure_PrintImage of a young sub adult male macaque (Macaca fascicularis unbrosus) observing an adult macaque using a pounding technique for breaking mature coconut in Great Nicobar Island, the mechanics of which are studied in this issue by Mazumder and Kaburu, on pages 141–159. Credit by Jayashree Mazumder.
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December 2019, Volume 40, Issue 6
10764_40_6_CoverFigure_PrintIllustration of an adult male golden-faced saki (Pithecia chrysocephala) of which the vocal repertoire is studied in this issue by Muir et al. on pages 721–743. Illustration by Jen Muir. Right-click here to enlarge the image




October 2019, Volume 40, Issue 4-5
10764_40_4_CoverFigure_PrintThis image represents an individual of black-fronted titi monkey (Callicebus nigrifrons) using the ground to consume fruit on the ground. This behavior was widely studied by Souza-Alves et al. on pages 553–572. Photo by Mariana Nagy-Reis.
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June 2019, Volume 40, Issue 3
10764_40_3_CoverFigure_PrintPhotograph of a juvenile François’ Langur (Trachypithecus francoisi) climbing on a shrub in a human-modified habitat at Mayanghe national nature reserve, China. Local attitudes toward langurs are studied to promote the sustainable conservation of this species in this issue by Niu et al. on pages 331–355. Photograph by Kefeng Niu.
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April 2019, Volume 40, Issue 2
10764_40_2_CoverFigure_PrintPhotograph of Gorilla gorilla gorilla “Mandara” at Smithsonian's National Zoo, Washington DC. “Mandara" is a participant in a study of orangutan and gorilla communication published in this issue by Botting and Bastian on pages 244–262. Photograph by Jennifer Botting.
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February 2019, Volume 40, Issue 1
10764_40_1_CoverFigure_PrintIllustration of the different primate hybridizations discussed in this Special Issue. [Lower left] T. crepusculus female. Photo credit: Tilo Nadler. [Lower right] Hybrid howler monkey (Alouatta palliata x A. pigra) in Tabasco, Mexico. Photo credit: Milagros González. [Upper left] Hybrid tamarin (Saguinus leucopus x S. geoffroyi) from Colombia. Photo credit: Iván Darío Soto Calderón. [Upper right] "UMA" a hybrid guenon (Cercopithecus mitis doggetti x C. ascanius schmidti) at Gombe National Park, Tanzania. Photo credit: Sarah Crawford, Gombe Hybrid Monkey Project.
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